I suspect it has more to do with the speed and wind resistance as they are aerodynamically efficient
I used to do a regular run of nearly 200 k on the Hume from Melbourne to Nagambie and always used 0.2 to 0.3 L/100 more going to nagambie and this is because of the change in elevation I suspect as there are more uphills than downhills.
But more noticeable was that if I sat on the speed limit of 110 compared to sometimes taking a more relaxed drive at say 100 there was a 1 to 1.5 l/100 difference.
For a while they had extensive road wire rope works and at 80 reduced speed for a considerable length saw the economy improve by a further 1 l/100 or so.
Wind resistance increases by a factor of 4 compared to speed .
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